喬雨軒 申瀟瀟 焦雪輝 周小娟 岳長(zhǎng)平 史喜兵
DOI:10.13925/j.cnki.gsxb.20230337
摘? ? 要:【目的】建立桃種質(zhì)的觀賞性綜合評(píng)價(jià)體系,為豫中地區(qū)合理引進(jìn)和應(yīng)用觀賞桃種質(zhì)資源提供參考依據(jù)?!痉椒ā繉?duì)46份桃種質(zhì)的花期長(zhǎng)度及其樹型、花色、花型、花徑、單枝著花數(shù)等性狀指標(biāo)進(jìn)行調(diào)查,采用層次分析法構(gòu)建模型,確定指標(biāo)權(quán)重,綜合評(píng)價(jià)其觀賞性?!窘Y(jié)果】在11個(gè)評(píng)價(jià)指標(biāo)中,花期長(zhǎng)度、花色、單枝著花數(shù)、花型、花徑所占權(quán)重比例均較大(>12%),是影響桃種質(zhì)觀賞價(jià)值的重要指標(biāo)。在賦分標(biāo)準(zhǔn)中,開張樹型、牡丹花型、紅花、大花徑、高花量、長(zhǎng)花期分值較高,均為5分。46份桃種質(zhì)花色繁多,主色以紅、粉、白為基調(diào),并有少量復(fù)色花種質(zhì);植株在豫中地區(qū)始花期集中于2月下旬至3月中下旬,嫣粉嬌香花期最長(zhǎng),為28 d,而緋桃和玲瓏粉花期最短,只有15 d;嫣紅早花的單枝著花數(shù)最高,平均為48.33朵;黃金美麗的平均花徑最大,為53.41 mm,且其花絲數(shù)也最多,平均達(dá)到89.67條;萬重粉的花瓣數(shù)是其他種質(zhì)的2~5倍,平均高達(dá)103.33枚。根據(jù)綜合得分,將46份桃種質(zhì)劃分為4個(gè)等級(jí),其中Ⅰ級(jí)7份,Ⅱ級(jí)17份,Ⅲ級(jí)11份,Ⅳ級(jí)11份?!窘Y(jié)論】利用層次分析法對(duì)桃種質(zhì)的觀賞性進(jìn)行評(píng)價(jià),較為系統(tǒng)全面、準(zhǔn)確有效。鄢紅、嫣紅早花、紅菊花桃、滿天紅、五寶桃、萬重粉和畫春壽星7份種質(zhì)得分均高于3.677 1,觀賞價(jià)值較高、適應(yīng)性強(qiáng),是園林應(yīng)用和雜交育種的優(yōu)良植物材料。
關(guān)鍵詞:桃種質(zhì);觀賞性狀;綜合評(píng)價(jià);層次分析法
中圖分類號(hào):S662.1 文獻(xiàn)標(biāo)志碼:A 文章編號(hào):1009-9980(2024)02-0216-13
Comprehensive evaluation of the ornamental value of 46 peach varieties in the central Henan
QIAO Yuxuan1, SHEN Xiaoxiao1, JIAO Xuehui1, ZHOU Xiaojuan1, YUE Changping2, 3*, SHI Xibing1*
(1Zhengzhou Institute of Agricultural and Technology Science, Zhengzhou 450005, Henan, China; 2Yanling Institute of Forestry Science, Ningling 461200, Henan, China; 3Yanling Donghua Planting Farmers Professional Cooperative, Ningling 461200, Henan, China)
Abstract:【Objective】 By screening traits closely related to ornamental value of peach germplasm, an ornamental evaluation system was constructed. Using this evaluation system, 46 peach varieties in the resource nursery were evaluated, with the aim of selecting varieties with high ornamental value and strong adaptability in central Henan. 【Methods】 11 ornamental traits of the 46 peach varieties were investigated. The quality traits of flowers were determined by comparing the peach blossom DUS testing guidelines; the length of flowering period was calculated by observing the start and end flowering periods of different variety groups. The approximate chilling requirement of the peach varieties was obtained by consulting the meteorological data of Yanling County from February to April on the China Meteorological Data Network and combining expert evaluations. The diameter of flowers in the vertical direction at full bloom was measured using vernier calipers. The number of flowers per branch was randomly selected from the upper 60 cm of the plant (20 cm for dwarf varieties), and the number of flowers on the branch was recorded. For the quantitative traits measured in various varieties, six consistent samples were selected for biological replicates and the average value was obtained. A questionnaire survey was used to determine the evaluation indicators and scoring criteria. An ornamental evaluation system for peach variety was constructed using the Analytical Hierarchy Process, consisting of a goal layer, a criteria layer, and a scheme layer. The consistency of the judgment matrix was tested using Yaahp software, and the weights of each indicator were calculated. Finally, the comprehensive score of ornamental quality of each peach variety was calculated. 【Results】Through pairwise comparisons of evaluation indicators, four judgment matrices were constructed, including A-C and C-P. Their consistency was tested and found to meet the requirements. In the criteria layer, the weight value of the flower quantity feature was the largest. Followed by the flower quality feature and the overall external feature. Among the 11 evaluation indicators in the solution layer, the length of flowering period, flower color, number of flowers per branch, flower shape, and flower diameter all had a large weight value (>12%), which are the important indicators affecting the ornamental value of peach germplasm. In the flower quality feature, the plumper flower shape, the more beautiful flower color, and the more prominent anther, the higher the score. All the flower quality indicators had a positive impact on the ornamental value of peach germplasm. Therefore, the higher the indicator level, the higher the score. There were many flower colors in the 46 peach varieties. The main colors included red, pink and white, with a small numbers of varieties with multicolor flowers. The plant flowering period in central Henan was mostly concentrated from late February to mid-to-late March. Among them, Yanfenjiaoxiang had the longest flowering period of 28 days; while Feitao and Linglongfen had the shortest flowering period of only 15 days. The cold requirement of different peach varieties varied greatly. In the planning and design of garden landscapes, the combination of different varieties of cold requirement could extend the peach ornamental period to about 50 days. Yanhongzaohua had the highest average number of flowers per branch, with an average of 48.33 flowers. When in full bloom, the flowers were clustered together, colorful, and the entire branch was covered with flowers. Huangjinmeili had the largest average flower diameter, reaching 53.41 mm. It also had the highest number of filaments, with an average of 89.67. This led to the orange anther being particularly eye-catching, with distinctive characteristics and a unique style. Wanchongfen had a significantly difference in number of petals from the other varieties, with an average of 103.33, which was 2-5 times that of ordinary ornamental peaches. It also had a large flower diameter and a peony-type flower shape, which was very unique and novel. According to the comprehensive score, the 46 peach varieties were clustered using the Euclid distance. The results were divided into four grades, including 7 at grade Ⅰ, 17 at grade Ⅱ, 11 at grade Ⅲ, and 11 at grade Ⅳ. 【Conclusion】The use of Analytic Hierarchy Process to evaluate the ornamental value of peach germplasm is relatively systematic, comprehensive, accurate and effective. Yanhong, Yanhongzaohua, Hongjuhuatao, Mantianhong, Wubaotao, Wanchongfen, and Huachunshouxing had scores >3.677 1, indicating high ornamental value and strong adaptability. These were excellent plant materials for landscape application and hybrid breeding. While Wanbai and Fenhongnvhai had low ornamental values and should be reduced in the central Henan. The other varieties with good ornamental value can be selected and utilized based on breeding objectives and application scenarios, combined with their own characteristics.
Key words: Peach germplasm; Ornamental character; Comprehensive evaluation; Analytical hierarchy process (AHP)
桃(Prunus persica L.)為薔薇科李屬桃亞屬植物,起源于中國(guó),栽培歷史達(dá)上千年,依果實(shí)品質(zhì)和花的觀賞特性,可將桃分為果桃和觀賞桃兩大類。其中觀賞桃具有古老的中國(guó)文化特色,成為文人墨客筆下美好的象征,是中國(guó)重要的早春觀花樹種[1]。因其耐旱耐寒、栽培難度低,且樹形多姿、花色豐富、花型豐腴,受到人們的廣泛喜愛,不僅在城市綠化、景區(qū)設(shè)計(jì)、專類園的建設(shè)中占有重要地位,而且對(duì)生態(tài)環(huán)境的改善也起到重要作用[2]?;ǖ奶匦允翘覙溆^賞性的集中表達(dá),針對(duì)其觀賞性狀的評(píng)價(jià)研究對(duì)優(yōu)良種質(zhì)的創(chuàng)制和景觀配置的利用具有重要的參考意義。
從現(xiàn)有資料來看,關(guān)于觀賞桃的研究主要集中于新品種培育[3-5]、遺傳多樣性分析[6-7]、分子標(biāo)記及基因功能驗(yàn)證[8-10]、生理生化分析[11-13]、種質(zhì)資源篩選利用[14-15]等方面,而對(duì)于花性狀的觀賞性評(píng)價(jià)研究則較少。劉衛(wèi)東等[16]對(duì)20個(gè)觀賞桃品種的引種適應(yīng)性進(jìn)行了綜合評(píng)價(jià),指標(biāo)的選取并未涉及觀賞特性。王立如等[17]僅對(duì)觀賞桃的生長(zhǎng)特性和花葉果性狀進(jìn)行了調(diào)查,在觀賞價(jià)值方面并未進(jìn)行系統(tǒng)評(píng)價(jià)。王力榮等[18]對(duì)30個(gè)桃品種的18個(gè)觀賞性狀進(jìn)行了調(diào)查分析,全面梳理了不同品種的觀賞特性并進(jìn)行綜合描述。對(duì)比前人研究,目前對(duì)桃種質(zhì)的觀賞性分析中大多未引入系統(tǒng)的評(píng)價(jià)體系,無法得到量化的數(shù)據(jù)結(jié)果和評(píng)價(jià)等級(jí)。層次分析法(analytical hierarchy process,AHP)作為一種定性與定量結(jié)合、系統(tǒng)化、層次化的分析方法,可將復(fù)雜問題進(jìn)行分解,適用于無法完全定量分析解決的難題,是一種將個(gè)人主觀判斷進(jìn)行定量化客觀描述的手段[19]。目前已成功應(yīng)用于梅花[20]、櫻花[21]、鳳仙花[22]、木蘭[23]、酢漿草[24]等多種植物的觀賞性綜合評(píng)價(jià)。迄今為止,針對(duì)桃的分析評(píng)價(jià)多基于花葉果的農(nóng)藝學(xué)性狀、生長(zhǎng)特性和適應(yīng)性,而針對(duì)花的觀賞特性運(yùn)用層次分析法構(gòu)建綜合評(píng)價(jià)體系的研究還未見報(bào)道,從而影響優(yōu)質(zhì)觀賞桃品種的推廣及應(yīng)用。
筆者在前期觀測(cè)統(tǒng)計(jì)資源圃桃屬植物的41個(gè)DUS測(cè)試性狀[25]的基礎(chǔ)上,運(yùn)用層次分析法,篩選出11個(gè)與桃種質(zhì)觀賞性密切相關(guān)的性狀指標(biāo)。從桃花的觀賞特性出發(fā),構(gòu)建出桃種質(zhì)的觀賞性評(píng)價(jià)體系,利用該評(píng)價(jià)體系對(duì)資源圃中46份桃種質(zhì)進(jìn)行評(píng)價(jià),以期篩選出觀賞性佳、適應(yīng)性強(qiáng)的種質(zhì)資源,為今后觀賞桃優(yōu)良種質(zhì)的創(chuàng)制及豫中地區(qū)桃花景觀的規(guī)劃設(shè)計(jì)提供科學(xué)依據(jù)。
1 材料和方法
1.1 試驗(yàn)材料
試驗(yàn)地位于河南省許昌市鄢陵縣林業(yè)科學(xué)研究所資源圃,北緯34°00′29″,東經(jīng)114°14′47″,海拔56 m,屬亞溫帶季風(fēng)性氣候,四季分明,年均氣溫為14.3 ℃,年均無霜期222 d,年均日照時(shí)數(shù)為2322 h,年均降水量為608.2 mm[26]。最熱月為7月,平均氣溫為27.3 ℃;最冷月在1月,平均氣溫為0.25 ℃。
本研究中選用的46份桃種質(zhì)的來源主要分為兩類,一類引種于中國(guó)農(nóng)業(yè)科學(xué)院鄭州果樹研究所,另一類來自鄢陵縣林業(yè)科學(xué)研究所選育的優(yōu)良雜交后代和長(zhǎng)期保存的種質(zhì)資源,其中開張形30個(gè),垂枝形5個(gè),龍柱形6個(gè),矮化形5個(gè)(表1),立地條件相當(dāng)、管理模式相同,觀察發(fā)現(xiàn)46份種質(zhì)生長(zhǎng)狀況良好,適應(yīng)性強(qiáng),未發(fā)生明顯的病蟲害。
1.2 試驗(yàn)方法
1.2.1 評(píng)價(jià)指標(biāo)的篩選 本試驗(yàn)調(diào)查問卷共2種。問卷一為專家問卷,運(yùn)用AHP法從外部整體特征、花質(zhì)量特征和花數(shù)量特征3個(gè)角度入手,請(qǐng)有多年觀賞桃栽培經(jīng)驗(yàn)的專家對(duì)41個(gè)DUS測(cè)試指標(biāo)進(jìn)行篩選和確定,并對(duì)評(píng)價(jià)指標(biāo)的重要性采用1~9標(biāo)度法[27]進(jìn)行兩兩比較。問卷二是收集被調(diào)查者(觀賞園藝相關(guān)從業(yè)者、愛好者)依主觀感受對(duì)各評(píng)價(jià)指標(biāo)子選項(xiàng)的意見,采用5分制構(gòu)建出桃種質(zhì)觀賞性評(píng)價(jià)的評(píng)分標(biāo)準(zhǔn),最終根據(jù)AHP法得到的指標(biāo)權(quán)重確定種質(zhì)的綜合得分。
問卷一于2023年5月22—29日共發(fā)放30份,收回30份且均為有效問卷;問卷二于2023年6月5-30日共發(fā)放110份,收回109份,其中有效問卷105份。
1.2.2 觀賞性狀調(diào)查與統(tǒng)計(jì) 2023年2月中旬至4月底,調(diào)查46份桃種質(zhì)的樹型、花型、花色、花期長(zhǎng)度、花徑、單枝著花數(shù)等觀賞性指標(biāo),其中花質(zhì)量性狀比對(duì)桃花DUS測(cè)試指南進(jìn)行確定;花期長(zhǎng)度通過觀測(cè)不同種質(zhì)群體的始花期(植株上5%花朵開放)和末花期(植株上75%花瓣變色)進(jìn)行計(jì)算,并通過中國(guó)氣象數(shù)據(jù)網(wǎng)查閱鄢陵縣2—4月氣象數(shù)據(jù),結(jié)合專家評(píng)估得出桃種質(zhì)的大致需冷量[28];使用游標(biāo)卡尺測(cè)量成熟盛開時(shí)垂直方向的花朵直徑;單枝著花數(shù)即隨機(jī)選取植株中上部60 cm長(zhǎng)的枝條(矮化型種質(zhì)選取20 cm),記錄枝條上著生的花朵數(shù)量。各種質(zhì)所測(cè)數(shù)量性狀均選取6個(gè)較一致樣本進(jìn)行生物學(xué)重復(fù)后取平均值,減少系統(tǒng)誤差。
1.2.3 AHP模型構(gòu)建及檢驗(yàn) 根據(jù)問卷一數(shù)據(jù)并結(jié)合專家意見,構(gòu)建由目標(biāo)層(A)、準(zhǔn)則層(C)、方案層(P)組成的桃種質(zhì)觀賞性評(píng)價(jià)體系。準(zhǔn)則層是確定評(píng)價(jià)的性狀類別,從桃的主要觀賞特性出發(fā),由于先花后葉的生長(zhǎng)習(xí)性,花器是桃種質(zhì)的主要觀賞部位,而樹型、生長(zhǎng)勢(shì)和香氣也是影響觀賞價(jià)值的重要因素,故將三者劃為一類,因此確定3個(gè)主要性狀為外部整體特征(C1)、花質(zhì)量特征(C2)、花數(shù)量特征(C3);方案層作為細(xì)化的二級(jí)指標(biāo)分為11個(gè)具體的評(píng)價(jià)因子(表2)。利用yaahp軟件對(duì)判斷矩陣的一致性進(jìn)行檢驗(yàn)并計(jì)算各指標(biāo)權(quán)重。當(dāng)一致性比率CR<0.10時(shí),矩陣一致性較高。
2 結(jié)果與分析
2.1 各層指標(biāo)的判斷矩陣
通過問卷一中對(duì)評(píng)價(jià)指標(biāo)的兩兩比較,構(gòu)建A~C、C~P共4個(gè)判斷矩陣,經(jīng)檢驗(yàn)一致性均符合要求(表3)。在準(zhǔn)則層(C)中,花數(shù)量特征(C3)的權(quán)重值最大(0.546 7),其次為花質(zhì)量特征(0.315 4)和外部整體特征(0.137 9),表明花數(shù)量特征是桃種質(zhì)觀賞性評(píng)價(jià)中最重要的性狀。在外部整體特征中,樹型所占權(quán)重最大(0.429 9),其次為生長(zhǎng)勢(shì)(0.400 8)和香氣(0.169 3)。在花質(zhì)量特征中,花色和花型占主導(dǎo)地位,權(quán)重值分別為0.487 7和0.444 6,而花藥顏色僅占比0.067 7。在花數(shù)量特征中,按貢獻(xiàn)比例大小排序依次為花期長(zhǎng)度、單枝著花數(shù)、花徑、花瓣數(shù)和花絲數(shù),其權(quán)重值分別為0.357 1、0.274 0、0.231 3、0.102 6和0.034 9。因此,花期長(zhǎng)度、單枝著花數(shù)、花徑、花色、花型是桃種質(zhì)觀賞性評(píng)價(jià)的關(guān)鍵指標(biāo)。
2.2 各指標(biāo)排名及賦分標(biāo)準(zhǔn)
由圖1可知,花期長(zhǎng)度對(duì)桃種質(zhì)的觀賞價(jià)值影響最大,其權(quán)重所占比例達(dá)19.52%,這與目前觀賞桃種質(zhì)的花期長(zhǎng)度差異較大且大多偏短有關(guān);其次花色、單枝著花數(shù)、花型和花徑的權(quán)重也均處于較高水平,分別為15.38%、14.98%、14.02%和12.65%,位列第2~5名;樹型、花瓣數(shù)、生長(zhǎng)勢(shì)的權(quán)重較為接近,分別為5.93%、5.61%、5.53%;而香氣、花藥顏色和花絲數(shù)則排名靠后,其所占權(quán)重分別為2.33%、2.13%和1.91%。
將46份桃種質(zhì)盛花期時(shí)的整體植株、花枝、單花照片制作幻燈片,根據(jù)問卷二調(diào)查結(jié)果,綜合專家意見和大眾喜好,構(gòu)建出5分制的桃種質(zhì)觀賞性評(píng)價(jià)賦分標(biāo)準(zhǔn)(表4)。在樹型中,由于開張型和垂枝型桃花在園林綠化中應(yīng)用較多,公眾和市場(chǎng)認(rèn)可度較高,因此分值相對(duì)較高。在花質(zhì)量特征中,花型越飽滿、花色越艷麗、花藥越突顯,則分值越高。而在花數(shù)量特征中,所有指標(biāo)均對(duì)桃種質(zhì)的觀賞價(jià)值呈正向影響,故指標(biāo)量級(jí)越高則分值越高。
2.3 桃種質(zhì)的質(zhì)量性狀、香味程度及需冷量
胡東燕等[29]以花型作為次級(jí)分類標(biāo)準(zhǔn),將觀賞桃的花型分為單瓣型、梅花型、月季型、菊花型和牡丹型。由表5可知,在調(diào)查的46份桃種質(zhì)中,依花型出現(xiàn)頻次從高到低依次為梅花型、月季型、單瓣型、菊花型和牡丹型,種質(zhì)數(shù)分別為17、16、5、4和4份。其中單瓣型的種質(zhì)為帚形山桃、山桃、毛桃、垂枝甘肅桃、甘肅桃,均為整株保存的原始種質(zhì)資源,而其余4種花型的種質(zhì)大多為培育種或選育種,說明花型的飽滿程度能提升桃花的觀賞價(jià)值,目前牡丹型種質(zhì)資源較少,仍是今后觀賞桃育種工作的重要方向。觀賞桃花色繁多,從白到粉,由粉至紅,單色轉(zhuǎn)復(fù)色,色彩繽紛,其主調(diào)以紅、粉、白三色為主,同一色系的不同種質(zhì)間花色深淺略有差別。在46份種質(zhì)中,單色花共39份,復(fù)色花7份,分別為含紅、灑紅、灑紅龍柱、灑紅垂枝、鴛鴦垂枝、人面和五寶桃,依復(fù)色部位和類型又可分為三類:第一類是灑紅、灑紅龍柱、灑紅垂枝和五寶桃,其植株就單一花朵來說既有單色花也有復(fù)色花。第二類是含紅、人面,其復(fù)色類型為兩種單色花,且兩種花可在同一花枝共生。第三類是鴛鴦垂枝,其復(fù)色類型與第二類相同,但復(fù)色部位極為規(guī)整,同一花枝的花色相同。從表5還可以看出,在豫中地區(qū)不同桃種質(zhì)的需冷量差異較大,低至毛桃的100 h,高至紅菊花桃和五寶桃的1250 h,需冷量跨度大閾值廣,為桃花景觀的配置利用提供了巨大空間。需冷量低的早花品種,如鄢紅、嫣紅早花、嫣碧蕊香等,2月底即進(jìn)入始花期,可極大彌補(bǔ)晚冬觀花的空缺,滿足大眾對(duì)早春賞桃花雪的熱烈需求;需冷量高的晚花品種,如滿天紅、人面、菊花桃等,始花期為3月末,可作為延長(zhǎng)觀花期的優(yōu)良配套品種,在各類園林景觀規(guī)劃設(shè)計(jì)中,合理運(yùn)用不同品種的需冷量進(jìn)行組合搭配,可將桃花觀賞期延長(zhǎng)至50 d左右。經(jīng)調(diào)查多數(shù)桃種質(zhì)均無香味,具有淡香的種質(zhì)共13份,主要包含探春、報(bào)春、灑紅、迎春及山桃、甘肅桃等原始種,而嫣粉嬌香、垂枝甘肅桃、嫣碧花香等5份種質(zhì)具有較濃烈的芳香味。
2.4 桃種質(zhì)的數(shù)量性狀
花的數(shù)量性狀是影響桃種質(zhì)觀賞價(jià)值的關(guān)鍵因素。從表6可以看出,單枝著花數(shù)超過40朵的種質(zhì)有6份,分別為嫣紅早花(48.33朵)、鄢紅(45.67朵)、垂枝甘肅桃(44.5朵)、紅線菊(43朵)、滿天紅(41朵)、山桃(40.67朵),其中嫣紅早花、鄢紅和滿天紅的花徑也較大,分別達(dá)到了43.06 mm、41.66 mm和47.1 mm,盛開時(shí)花團(tuán)錦簇,色彩艷麗,整個(gè)枝條布滿花朵,花量和花徑的兼顧性較高,而紅線菊雖花徑僅有34.89 mm,但花型特殊,同樣具有較高的觀賞價(jià)值。黃金美麗的平均花徑最大,為53.41 mm,且花絲數(shù)也最多,平均達(dá)到89.67條,導(dǎo)致橘色花藥格外醒目,特點(diǎn)突出,別具一格。從花瓣數(shù)來看,萬重粉明顯有別于其他種質(zhì),平均高達(dá)103.33枚,是普通觀賞桃的2~5倍,且花徑大、花型為牡丹型,十分獨(dú)特,新穎別致。參試材料的花期長(zhǎng)度大多在17~20 d,花期超過22 d的種質(zhì)有6份,分別為嫣粉嬌香、鄢紅、嫣紅早花、畫春壽星、探春、粉垂菊,其中嫣粉嬌香開花持續(xù)時(shí)間最久,長(zhǎng)達(dá)28 d,以上種質(zhì)均可作為彌補(bǔ)桃花花期短的優(yōu)良種質(zhì)。
2.5 桃種質(zhì)綜合得分及等級(jí)分類
按照2.2中制定的賦分標(biāo)準(zhǔn),對(duì)46份桃種質(zhì)各指標(biāo)的觀測(cè)結(jié)果進(jìn)行賦值,參照權(quán)重將加權(quán)后的各指標(biāo)分值相加得到桃種質(zhì)的觀賞性綜合得分(表7)。根據(jù)綜合得分,采用歐氏距離進(jìn)行聚類,將46份桃種質(zhì)劃分為4個(gè)等級(jí)(圖2),其中Ⅰ級(jí)(>3.677 1),7份;Ⅱ級(jí)(3.216 4~3.677 1),17份;Ⅲ級(jí)分值(2.856 0~3.085 9),11份;Ⅳ級(jí)分值(<2.856 0),11份。
其中,Ⅰ級(jí)的鄢紅、嫣紅早花、紅菊花桃、滿天紅等7份種質(zhì),均具有花色艷麗、花徑大、單枝著花數(shù)多、花期較長(zhǎng)的特點(diǎn),說明花色、花徑、花期長(zhǎng)度、單枝著花數(shù)是衡量桃種質(zhì)觀賞性的重要指標(biāo),可將其作為觀花型桃屬植物培育及篩選應(yīng)用的參考依據(jù)。紅線菊和粉垂菊在Ⅱ級(jí)中脫穎而出,僅此二者的瓣形為單瓣且花徑較小,其他種質(zhì)均為復(fù)瓣或重瓣,說明僅憑花徑、花瓣數(shù)量并不能決定桃種質(zhì)的觀賞價(jià)值,相比之下,獨(dú)特的花型、較長(zhǎng)的花期、高單枝著花數(shù)顯得更為重要。另外,大部分桃屬植物的花香很淡甚至沒有,而具有較濃花香的探春、嫣粉嬌香這2份種質(zhì),在香氣指標(biāo)上得分最高,導(dǎo)致綜合排名也較高。因此,香氣這一指標(biāo)可作為觀花型桃屬植物選育的特殊參考因素和研究方向。Ⅲ級(jí)和Ⅳ級(jí)主要分布在龍柱桃和原始種,其中龍柱桃大多存在花期較短、單枝著花數(shù)少的問題,盛開時(shí)略顯稀疏單薄,因此如何延長(zhǎng)花期并增加花量是龍柱桃新品種培育中亟待解決的問題。原始種雖不適合園林應(yīng)用,但包含了豐富多樣的遺傳信息,是優(yōu)良的種質(zhì)資源,其育種價(jià)值遠(yuǎn)大于觀賞價(jià)值。
3 討 論
園林植物的觀賞性評(píng)價(jià)方法還有許多,例如模糊評(píng)價(jià)法、灰色關(guān)聯(lián)法、百分制計(jì)分法[30-32]等。在對(duì)桃種質(zhì)進(jìn)行觀賞性評(píng)價(jià)時(shí),由于樣本群體大、評(píng)價(jià)指標(biāo)多且關(guān)聯(lián)度高,既有質(zhì)量性狀,又有數(shù)量性狀,因此,選用AHP法能實(shí)現(xiàn)定性與定量的統(tǒng)一轉(zhuǎn)化,確保評(píng)價(jià)結(jié)果的科學(xué)性和有效性。AHP法的核心內(nèi)容為層次結(jié)構(gòu)的建立和指標(biāo)權(quán)重的確定,禹霖等[33]在運(yùn)用AHP法對(duì)18個(gè)觀賞桃品種的評(píng)價(jià)研究中,未建立層級(jí)結(jié)構(gòu)并默認(rèn)各指標(biāo)因子權(quán)重相當(dāng),方法缺乏科學(xué)性,因此評(píng)價(jià)結(jié)果有失偏頗,參考價(jià)值有待商榷。另外,在以往大多數(shù)植物評(píng)價(jià)研究中,對(duì)方案層指標(biāo)的量化打分,多以征詢專家意見并結(jié)合參考文獻(xiàn)的方式制定賦分標(biāo)準(zhǔn)。為針對(duì)桃種質(zhì)制定一套最貼近大眾日常審美的評(píng)分標(biāo)準(zhǔn),筆者在本研究中采用幻燈片結(jié)合問卷的形式,讓受訪者根據(jù)自身喜好對(duì)各指標(biāo)的子選項(xiàng)進(jìn)行排序,根據(jù)最終排序結(jié)果確定分值。該方法最大限度地避免了個(gè)人因素的主觀性和偶然性,有助于確保評(píng)價(jià)結(jié)果的合理性。
觀賞桃作為極具開發(fā)價(jià)值和應(yīng)用前景的春花樹種,對(duì)其觀賞價(jià)值進(jìn)行評(píng)價(jià)是極為重要的基礎(chǔ)性工作,可為桃花品種的引種選擇、開發(fā)利用、雜交親本篩選及新品種培育等提供重要參考和科學(xué)依據(jù)。目前,對(duì)觀賞桃的評(píng)價(jià)研究多集中于園林應(yīng)用方面,對(duì)矮化型桃的綜合評(píng)價(jià)鮮有報(bào)道。如王立如等[17]對(duì)來自中國(guó)農(nóng)業(yè)科學(xué)院鄭州果樹研究所和江蘇農(nóng)業(yè)科學(xué)院的18個(gè)觀賞桃品種,開展了物候期、花葉果性狀、生長(zhǎng)特性等的觀測(cè)調(diào)查,研究對(duì)象均為開張型、垂枝型或龍柱型觀賞桃品種,更傾向于城市園林的應(yīng)用。筆者在本研究中加入了5份矮化型桃種質(zhì),但由于矮化型桃更適于盆栽觀賞,與園林應(yīng)用上的長(zhǎng)枝型桃花在觀賞方式和觀賞價(jià)值上存在差異,相比而言矮化型桃更強(qiáng)調(diào)對(duì)個(gè)體的展示,而長(zhǎng)枝型觀賞桃則以園林觀賞價(jià)值為基礎(chǔ),側(cè)重群體效果的表達(dá),這也是在綜合得分中,半矮化型的畫春壽星得分較高,而其余4份矮化型桃得分普遍較低的原因。因此將矮化型桃作為單獨(dú)一類進(jìn)行觀賞性評(píng)價(jià)會(huì)更加準(zhǔn)確合理,在評(píng)價(jià)指標(biāo)和評(píng)分標(biāo)準(zhǔn)的制定上應(yīng)更具針對(duì)性。另外,筆者在調(diào)查觀賞桃花色時(shí)發(fā)現(xiàn),桃花色彩繁多,紅色和粉色由淺及深,各不相同,除單色花外還有復(fù)色花,且不同種質(zhì)的復(fù)色類型和復(fù)色部位也不盡相同。筆者在本研究中針對(duì)花色指標(biāo)擬定的5級(jí)分類標(biāo)準(zhǔn),仍不足以全面地展示觀賞桃花色的多樣性,之后對(duì)花色還需開展更加細(xì)化的分類研究。
在準(zhǔn)則層3個(gè)指標(biāo)中,花數(shù)量特征所占權(quán)重最大,其次為花質(zhì)量特征,而外部整體特征所占權(quán)重最小,這與戴中武等[34]對(duì)獨(dú)蒜蘭屬植物觀賞性的判斷結(jié)果相同,并與花器是桃最主要觀賞部位的特性相符。從綜合得分來看,本試驗(yàn)位列Ⅰ級(jí)的桃種質(zhì)均具有花色艷麗、花徑大、花量多、花期較長(zhǎng)的特點(diǎn),這與董鈉等[35]的研究結(jié)果一致。在各評(píng)價(jià)指標(biāo)權(quán)重值排序上,對(duì)觀賞性貢獻(xiàn)率最高的為花期長(zhǎng)度,其次為花色、單枝著花數(shù)、花型、花徑、樹型、花瓣數(shù)、生長(zhǎng)勢(shì),而排名靠后的是香氣、花藥顏色和花絲數(shù)。由于不同樹型的桃種質(zhì)間差異較大,本研究中的矮化型桃評(píng)分較低與實(shí)際園林應(yīng)用中的情況吻合,隨后可針對(duì)適用于盆栽型的觀賞桃品種進(jìn)行評(píng)價(jià)篩選,對(duì)本文研究結(jié)果加以補(bǔ)充。為使篩選出的桃種質(zhì)能早日投入生產(chǎn)并推廣普及,實(shí)現(xiàn)科技興花,仍需深入研究。
4 結(jié) 論
本研究中,基于AHP法建立的桃種質(zhì)觀賞性評(píng)價(jià)模型共分為3個(gè)層次,包含了與桃種質(zhì)觀賞性密切相關(guān)的11個(gè)指標(biāo)因子,較為系統(tǒng)全面。在11個(gè)指標(biāo)中,花期長(zhǎng)度、花色、單枝著花數(shù)、花型、花徑所占權(quán)重較大,是影響桃種質(zhì)觀賞性最重要的因素,可作為觀賞桃園林應(yīng)用選擇時(shí)的主要依據(jù)。
46份桃種質(zhì)中鄢紅、嫣紅早花、紅菊花桃、滿天紅、五寶桃、萬重粉、畫春壽星的觀賞價(jià)值較高,是園林應(yīng)用和雜交育種的優(yōu)良植物材料;而晚白、粉紅女孩等觀賞價(jià)值較低,在豫中地區(qū)應(yīng)減少應(yīng)用;其他種質(zhì)觀賞性良好,可根據(jù)育種目標(biāo)和應(yīng)用場(chǎng)景,結(jié)合其自身特性加以選擇利用。
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收稿日期:2023-08-29 接受日期:2023-12-12
基金項(xiàng)目:中原學(xué)者工作站(ZYGZZ2022073)
作者簡(jiǎn)介:?jiǎn)逃贶?,男,研究?shí)習(xí)員,碩士,主要從事園林植物育種及應(yīng)用研究。E-mail:294694128@qq.com
*通信作者 Author for correspondence. E-mail:3511319300@qq.com;E-mail:madary@163.com